Just as a fish does not understand the importance of water while swimming in it but she realises its importance when separated from it and dies yearning for are-union.
Just as a deer and a bird living in a jungle do not realize its importance but realises its significance when caught and put in a cage by the hunter and wail for going back to the jungle.
Just as a wife does not appreciate the importance of staying with her husband when together but comes to her senses when separated from her husband. She wails and cries due to pangs of separation from him.
Similarly, a seeker living in the refuge of the True Guru remains oblivious of the greatness of Guru. But when separated from Him, repents and laments. (502)